Ilia State University

Status State University

Abbreviation ISU

Telephone (+995 32) 222 00 09



Address Kakutsa Cholokashvili Ave 3/5, , Tbilisi, Tbilisi, Georgia

Contact person Giga Zedania (Rector)

Contact person David Tarknishvili (Dean)

Departments Institute of Ecology , 4D Research Institute , Institute of Botany , Institute of Earth Sciences , Abastumani Astrophysical Observatory , The International School for Caucasus Studies (ISCS) , Giorgi Tsereteli Institute of Oriental Studies ,

Projects Promoting Sustainable Dam Development at River-Basin-Scale in the Southern Caucasus , Ethnobotany of the Caucasus , Investigation of Neotectonic deformation of the wester portion of Kura fold thrust belt ,

Academic programs Sustainable Natural and Forest Resources Management (MBA) at Ilia State University, Tbilisi, Georgia , Natural and Forest Resources and Biodiversity (MSC) at Ilia State University, Tbilisi , Ecology (ME) at Ilia State University, Tbilisi , International school for caucasus studies at Ilia State University, Tbilisi , BACHELOR IN BUSINESS ADMINISTRATION (BBA - MANAGEMENT, TOURISM) ,


Avtandil Okrostsvaridze

David Tarkhnishvili

Levan Mumladze

Jana Ekhvaia

Alexander Gavashelishvili

Marine Murtskhvaladze

Zaal Kikvidze

Nato Tephnadze

Natalia Togonidze

Tea Mumladze

Nugzar Zazanashvili

Maia Akhalkatsi

George Nakhutsrishvili

Bella Japoshvili

Nina Shatberashvili

Nikoloz Lachashvili

Mzia Kokhia

George Anchabadze

Goderdzi Didebulidze

Khatuna Gigauri

Natia Kopaliani

Nikoloz Tushabramishvili

Gigi Tevzadze

Elene Natroshvili

Mariam Gabelaia

Timothy Blauvelt

Mikheil Elashvili

Nino Skhirtladze

Zezva Asanidze

Zura Javakhishvili

Marine Mosulishvili

Nana Bakhtadze

Nato Jorjiashvili

Nana Gratiashvili

Giorgi Sokhadze

Ghia Nodia

Tengiz Verulava

Ketevan Gurchiani

Giorgi Boichenko

Tea Godoladze

Lasha Sukhishvili

Zurab Javakhishvili

Irakli Skhirtladze

Eter Abulidze

Ioseb Natradze

Rayya El Zein

Levan Ninua


Okrostsvaridze A., (2015) Ore occurences in the Georgian segment of the eastern greater caucasus: New research results. Bulletin of the Georgian National Academy of Sciences, vol: 9, 102-110

Tarkhnishvili D., (2012) Palaeoclimatic models help to understand current distribution of Caucasian forest species. Biological Journal of the Linnean Society, vol: 105, 231-248

Tarkhnishvili D., Gavashelishvili A., Murtskhvaladze M., Gabelaia M., Tevzadze G., (2014) Human paternal lineages, languages, and environment in the caucasus. Human Biology, vol: 86, 113-130

Mumladze L., (2008) A new species of the genus Helix from the lesser caucasus (SW Georgia). Journal of Conchology, vol: 39, 483-485

Gavashelishvili A., (2004) Habitat selection by East Caucasian tur (Capra cylindricornis). Biological Conservation, vol: 120, 395-402

Ekhvaia J., (2014) Genetic diversity of Vitis vinifera in Georgia: relationships between local cultivars and wild grapevine, V. vinifera L. subsp. sylvestris. Genetic Resources and Crop Evolution, vol: 61, 1507-1521

Callaway R., Kikvidze Z., (2000) Facilitation by unpalatable weeds may conserve plant diversity in overgrazed meadows in the Caucasus Mountains . Oikos, vol: 89, 275-282

Tarkhnishvili D., (2014) Historical biogeography of the caucasus. Nova Science Publishers, Inc., New York.

Tarkhnishvili D., (2008) Ecological divergence between two evolutionary lineages of the Caucasian salamander: Evidence from GIS analysis. Herpetological Journal, vol: 18, 155-163

Murtskhvaladze M., Gavashelishvili A., Tarkhnishvili D., (2010) Geographic and genetic boundaries of brown bear (Ursus arctos) population in the Caucasus. Molecular Ecology, vol: 19, 1829-1841

Gokhelashvili R., Javakhishvili Z., (2002) Tourism Infrastructure Planning in Tusheti National Park of Georgia. CONFERENCE PAPER: Conference proceedings of monitoring and management of visitor flows in recreational and protected areas, 74-77

Kikvidze Z., (2014) Loss of traditional knowledge aggravates wolf–human conflict in Georgia (Caucasus) in the wake of socio-economic change. Ambio, vol: 44, 452-457

Gavashelishvili A., (2010) Combining radio-telemetry and random observations to model the habitat of Near Threatened Caucasian grouse Tetrao mlokosiewiczi. ORYX, vol: 44, 491-500

Tephnadze N., et al. (2014) The impacts of management and site conditions on the phytodiversity of the upper montane and subalpine belts in the Central Greater Caucasus. Phytocoenologia, vol: 44, 255-282

Sayim F., Tarkhnishvili D., (2009) Some water chemistry parameters of breeding habitats of the Caucasian salamander, Mertensiella caucasica in the Western Lesser Caucasus. Comptes Rendus - Biologies, vol: 332, 464-469

Togonidze N., (2015) Variability of plant species diversity during the natural restoration of the subalpine birch forest in the Central Great Caucasus. Turkish Journal of Botany, vol: 39, 458-471

Mumladze T., Forte A., Cowgill E., et al. (2015) Subducted, detached, and torn slabs beneath the Greater Caucasus. GeoResJ, vol: 5, 36–46

Tarkhnishvili D., (2001) Humid forest refugia, speciation and secondary introgression between evolutionary lineages: Differentiation in a Near Eastern brown frog, Rana macrocnemis. Biological Journal of the Linnean Society, vol: 74, 141 - 156

Tarkhnishvili D., (2016) Mitochondrial phylogeny of the darevskia saxicola complex: Two highly deviant evolutionary lineages from the easternmost part of the range. Herpetological Journal, vol: 26, 175-182

Zazanashvili N., et al. (2011) Strategic Guidelines for Responding to Impacts of Global Climate Change on Forests in the Southern Caucasus (Armenia, Azerbaijan, Georgia). WWF,

Akhalkatsi M., (2015) Forest Habitat Restoration in Georgia, Caucasus Ecoregion. Mtsignobari, Georgia.

Zazanashvili N., (2006) An Ecoregional Conservation Plan For The Caucasus. WWF, Tbilisi.

Mumladze L., (2015) Oribatid mite communities along an elevational gradient in Sairme gorge (Caucasus). Experimental and Applied Acarology, vol: 66, 41-51

Nakhutsrishvili G., (2015) Colchic and Hyrcanian forests of the Caucasus: Similarities, differences and conservation status. Flora Mediterranea, vol: 25, 185-191

Gavashelishvili A., (2008) Modelling the habitat requirements of leopard Panthera pardus in west and central Asia. Journal of Applied Ecology, vol: 45, 579-588

Shatberashvili N., et al. (2015) Outlook on climate change adaptation in the South Caucasus mountains.

Akhalkatsi M., (2016) Landraces and Wild Species of the Secale Genus in the Georgia (Caucasus Ecoregion). Open Access Journal, vol: 1,

Gavashelishvili A., (2006) Planning the conservation of the breeding population of cinereous vultures Aegypius monachus in the Republic of Georgia. ORYX, vol: 40, 76-83

Japoshvili B., Mumladze L., (2013) Invasive Carassius carp in Georgia: Current state of knowledge and future perspectives. Current Zoology, vol: 59, 732-739

Japoshvili B., (2012) Long-term assessment of a vendace (Coregonus albula L.) stock in Lake Paravani, South Georgia. Advances in Limnology, vol: 63, 363–369

Japoshvili B., (2009) Monitoring on ichthyofauna in Paravani Lake.

Shatberashvili N., et al. (2016) The Georgian Road Map on Climate Change Adaptation.

Tarkhnishvili D., Murtskhvaladze M., Gavashelishvili A., (2013) Speciation in Caucasian lizards: Climatic dissimilarity of the habitats is more important than isolation time. Biological Journal of the Linnean Society, vol: 109, 876-892

Akhalkatsi M., Ekhvaia J., Nakhutsrishvili G., et al. (2010) Reasons and processes leading to the erosion of crop genetic diversity in mountainous regions of Georgia. Mountain Research and Development, vol: 30, 304-310

Akhalkatsi M., Nakhutsrishvili G., (2006) Facilitation of Seedling Microsites by Rhododendron Caucasicum Extends the Betula Litwinowii Alpine Treeline, Caucasus Mountains, Republic of Georgia. Arctic, Antarctic, and Alpine Research, vol: 38, 481-488

Akhalkatsi M., (1996) Reproductive phenology and seed development of Gentianella caucasea in different habitats in the Central Caucasus. Flora, vol: 191, 161-168

Akhalkatsi M., (2012) Diversity and Genetic Erosion of Ancient Crops and Wild Relatives of Agricultural Cultivars for Food: Implications for Nature Conservation in Georgia (Caucasus). Perspectives on Nature Conservation - Patterns, Pressures and Prospects, InTech, 51-92

Matcharashvili T., Chelidze T., Jorjiashvili N., (2011) Non-extensive statistical analysis of seismicity in the area of Javakheti, Georgia. Computers and Geosciences, vol: 37, 1627-1632

Tarkhnishvili D., Gavashelishvili A., et al. (2001) Land Use and Landscape Conservation in Southern Georgia.

Lachashvili N., (2015) New data on some rare species of flora of Georgia . Bulletin of the Georgian National Academy of Sciences, vol: 9, 135-138

Zazanashvili N., (2000) Main types of vegetation zonation on the mountains of the Caucasus. Acta Phytogeographica Suecica, vol: 85, 7-16

Kokhia M., (2010) High mountain meadows mesofauna for pastures rehabilitation.

Kikvidze Z., (1996) Neighbour Interaction and Stability in Subalpine Meadow Communities. Journal of Vegetation Science, vol: 7, 41-44

Nakhutsrishvili G., (1998) The Vegetation of the Subnival Belt of the Caucasus Mountains. INSTAAR, University of Colorado, vol: 30, 222-226

Nakhutsrishvili G., (2012) The Vegetation of Georgia (South Caucasus).

Anchabadze G., (2012) The North Caucasus factor in the Georgian-Abkhaz conflict context. International Alert 2012,

Didebulidze G., (2016) The inter-annual distribution of cloudless days and nights in Abastumani: Coupling with cosmic factors and climate change. Journal of Atmospheric and Solar-Terrestrial Physics, vol: 141, 48-55

Walther F., Mumladze L., et al. (2014) Biogeography of the Land Snails of the Caucasus Region. International Union for Conservation of Nature,

Nakhutsrishvili G., (1990) On survival strategies of plants in the subnivale belt of the Caucasus. Giornale Botanico Italiano, vol: 124, 781-791

Adamia S., Mumladze T., Tsereteli E., Tsereteli N., et al. (2008) Late Cenozoic Tectonics and Geodynamics of Georgia (SW Caucasus). Georgian International Journal of Science and Technology ISSN ,

Bussmann R., Kikvidze Z., et al. (2016) A comparative ethnobotany of Khevsureti, Samtskhe-Javakheti, Tusheti, Svaneti, and Racha-Lechkhumi, Republic of Georgia (Sakartvelo), Caucasus. Journal of Ethnobiology and Ethnomedicine, 12-43

Gigauri K., Akhalkatsi M., Nakhutsrishvili G., (2016) Alpine plant distribution and thermic vegetation indicator on Gloria summits in the Central Greater Caucasus. Pakistan Journal of Botany, vol: 48, 1893-1902

Gigauri K., Akhalkatsi M., Nakhutsrishvili G., (2013) Monitoring of vascular plant diversity in a changing climate in the alpine zone of the Central Greater Caucasus. Turkish Journal of Botany, vol: 37, 1104-1114

Bock J., Nakhutsrishvili G., (1986) International symposium on high mountain ecology, Tbilisi- Kazbegi, Georgia, USSR. Arctic & Alpine Research, vol: 18, 121-122

Akhalkatsi M., (2015) Erosion and Prevention of Crop Genetic Diversity Landraces of Georgia (South Caucasus). Genetic Diversity and Erosion in Plants, 159-187

Nakhutsrishvili G., (2014) Georgia. Red List of Endemic Plants of the Caucasus: Armenia, Azerbaidjan, Georgia, Iran, Russia, and Turkey, Missouri Botanical Garden Press, Tbilisi,

Box E., Nakhutsrishvili G., Zazanashvili N., et al. (2000) Vegetation and Landscapes of Georgia (Caucasus), as a Basis for Landscape Restoration. Bulletin of the Institute of Environmental Science and Technology, Yokohama National University, vol: 26, 69-102

Kopaliani N., Tarkhnishvili D., (2014) Gene Flow Between Wolf and Shepherd Dog Populations in Georgia (Caucasus). Journal of heredity, vol: 105, 345-353

Pilot M., Kopaliani N., et al. (2014) Genetic Variability of the Grey Wolf Canis lupus in the Caucasus in Comparison with Europe and the Middle East: Distinct or Intermediary Population?. PloS One, vol: 9, 1-10

Murtskhvaladze M., Kopaliani N., Tarkhnishvili D., (2012) Gene introgression between Gazella subgutturosa and G. marica: Limitations of maternal inheritance analysis for species identification with conservation purposes. Acta Theriologica, vol: 57, 383–386

Zazanashvili N., et al. (2009) Territorial protection of globally threatened species in the Caucasus. Status and Protection of Globally Threatened Species in the Caucasus, Contour Ltd, Tbilisi, 222-231

Mancheno C., Zazanashvili N., (2016) Effectiveness of the network of protected areas of the South Caucasus at representing terrestrial ecosystems after the dissolution of the Soviet Union. Environmental Conservation, 1-8

Kopaliani N., (2009) Status of turs in Georgia and conservation action plan. Status and Protection of Globally Threatened Species in the Caucasus, Contour Ltd, Tbilisi, 61-68

Yarovenko Y., Zazanashvili N., (2016) Recent hard evidence for the occurrence of the Leopard, Panthera pardus (Mammalia: Felidae), in the Eastern Greater Caucasus. Zoology in the Middle East, vol: 62, 88-90

Zazanashvili N., Tarkhnishvili D., (2016) CAUCASUS.

Nakhutsrishvili G., Akhalkatsi M., (2013) First signs of transformation of alpine vegetation In the Caucasus as a result of climate warming (on the example of Kazbegi region).

Zazanashvili N., (1991) Relationship between soil acidity, types of phytocoenosis and bedrocks in high mountains of the Central Caucasus (In Georgian). Bulletin of the Georgia Academy of Science ,

Zazanashvili N., (2005) The Caucasus Ecoregional Conservation Plan as a Tool for the Implementatio of CBD at the Regional Scale.

Zazanashvili N., et al. (2007) Conservation of the Leopard in the Caucasus. CAT News, 4-8

Breitenmoser U., Zazanashvili N., (2007) General conditions for the conservation of the leopard in the Caucasus. CAT News, 34-39

Gavashelishvili A., (2009) GIS-based Habitat Modeling of Mountain Ungulate species in the Caucasus Hotspot.

Bohn U., Zazanashvili N., Nakhutsrishvili G., (2007) The Map of the Natural Vegetation of Europe and its application in the Caucasus Ecoregion. BULLETIN OF THE GEORGIAN NATIONAL ACADEMY OF SCIENCES, vol: 1, 112-121

Tushabramishvili N., (2015) Current Middle & Upper Palaeolithic research in the southern Caucasus. Antiquity , 927-928

Tevzadze G., (2016) Ethno-ecological contexts of the Skhalta Gorge and the Upper Svaneti (Georgia, the Caucasus). Journal of Political Ecology, vol: 23, 246-262

Tevzadze G., (2014) Project Svaneti. Svanetian Towers and Svanetian high art. History and hypothesis. Kadmos, 1-6

Tevzadze G., (2009) The Birth of the Georgian Nation. Identity and Ideology. Political and Societal Identities. Nationality and Religiosity. Identity Studies, 1-17

Akhalkatsi M., Tarkhnishvili D., (2012) Habitats of Georgia.

Hille A., Tarkhnishvili D., (2002) Morphometric, biochemical and molecular traits in Caucasian wood mice (Apodemus/Sylvaemus), with remarks on species divergence. Acta theriologica, vol: 47, 389-416

Kikvidze Z., (1993) Plant species associations in alpine‐subnival vegetation patches in the Central Caucasus. Journal of Vegetation Science, vol: 4, 297-302

Natroshvili E., (2016) Sustainable development of Georgian region Svaneti.

Gavashelishvili A., (2012) Movements and habitat use by immature Cinereous Vultures (Aegypius monachus) from the Caucasus. Bird Study, vol: 59, 449-462

Gabelaia M., Tarkhnishvili D., (2017) Phylogenetic signals in scale shape in Caucasian rock lizards (Darevskia species). Zoologischer Anzeiger - A Journal of Comparative Zoology,

Bussmann R., Kikvidze Z., et al. (2016) Plant and fungal use in Tusheti, Khevsureti, and Pshavi, Sakartvelo (Republic of Georgia), Caucasus. Acta Societatis Botanicorum Poloniae , vol: 86, 1-45

Kvavadze E., Mumladze L., (2008) Botanical and zoological remains from an early medieval grave at Tsitsamuri, Georgia. Vegetation History and Archaeobotany, vol: 17, 217–224

Kikvidze Z., (2006 ) Seasonal shifts in competition and facilitation in subalpine plant communities of the central Caucasus. Journal of Vegetation Science, vol: 17, 77-82

Kikvidze Z., (2009 ) Facilitation and interference in subalpine meadows of the central Caucasus. Journal of Vegetation Science , vol: 12, 833-838

Kikvidze Z., (2005) Small-scale guild proportions and niche complementarity in a Caucasian subalpine hay meadow. Journal of Vegetation Science, vol: 16, 565 - 570

Kikvidze Z., (2003) Modelling species richness and diversity in grassland communities of the Central Caucasus. Oikos , vol: 89, 123 - 127

Erschbamer B., Akhalkatsi M., et al. (2010) Plant diversity along altitudinal gradients in the Central Alps (South Tyrol, Italy) and in the Central Greater Caucasus (Kazbegi region, Georgia). Tuexenia, vol: 30, 11–29

Akhalkatsi M., (1997) Comparative embryology of three Gentianaceae species from the Central Caucasus and the European Alps. Plant Systematics and Evolution, vol: 204, 39–48

Blauvelt T., (2017) Pedagogy, modernity and nationalism in the Caucasus in the age of reaction, 1880–1905. Caucasus Survey , vol: 5, 121-141

Blauvelt T., (2015) Resistance, Discourse and Nationalism in the March 1956 Events in Georgia - from Blauvelt and Smith, eds. Georgia after Stalin: Nationalism and Soviet Power (2015). Routledge, London.

Blauvelt T., (2017) Familiar Strangers: The Georgian Diaspora and the Evolution of Soviet Empire.

Blauvelt T., (2013) Endurance of the Soviet imperial tongue: the Russian language in contemporary Georgia. Central Asian Survey , vol: 32, 189-209


Blauvelt T., (2016) The Muslim uprising in Ajara and the Stalinist revolution in the periphery. Nationalities Papers The Journal of Nationalism and Ethnicity , vol: 44, 359-379

Akhalkatsi M., Togonidze N., Asanidze Z., et al. (2017) Agrobiodiversity and genetic erosion of crop varieties and plant resources in the Central Great Caucasus. Annals of Agrarian Science, vol: 15, 11–16

Lachashvili N., (2017) Conspectus of trees and shrubs of Tbilisi environs (East Georgia, South Caucasus). Annals of Agrarian Science, vol: 15, 118–129

Lachashvili N., (2017) Steppe of Tbilisi environs (East Georgia, South Caucasus). Annals of Agrarian Science,

Elashvili M., et al. (2011) Investigation of Active Fault Scarps by Means of Geophysical Prospecting Methods, Javakheti Fault Case, Georgia. American Geophysical Union,

Japoshvili B., (2016) A review of benthic fauna biodiversity in Georgia. Annals of Agrarian Science, vol: 14, 7-10

Skhirtladze N., (2016) Identity processes and statuses in post-Soviet Georgia: Exploration processes operate differently. Journal of Adolescence, vol: 47, 197-209

Asanidze Z., Akhalkatsi M., (2014) Genetic relationships between wild progenitor pear (Pyrus L.) species and local cultivars native to Georgia, South Caucasus. Flora - Morphology, Distribution, Functional Ecology of Plants, vol: 209, 504–512

Asanidze Z., Akhalkatsi M., (2011) Comparative morphometric study and relationships between the Caucasian species of wild pear (Pyrus spp.) and local cultivars in Georgia. Flora - Morphology, Distribution, Functional Ecology of Plants, vol: 206, 974-986

Mumladze L., Asanidze Z., Japoshvili G., (2017) An inverse elevational species richness gradient of Caucasian vascular plants and Encyrtidae (Hymenoptera, Chalcidoidea). Ecoscience, 1-5

Mumladze L., Asanidze Z., (2017) Beyond elevation: testing the climatic variability hypothesis vs. Rapoport’s rule in vascular plant and snail species in the Caucasus. Biological Journal of the Linnean Society, vol: 121, 753–763

Akhalkatsi M., Asanidze Z., (2015) Pyrus caucasica Fed. Ancestor of Georgian Pear Landraces. Science, Technology and Development, vol: 34, 105-108

Asanidze Z., Akhalkatsi M., (2011) Morphological Relationships between Wild and Cultivated Pears in Georgia. Proceedings of the Georgian Academy of Sciences, vol: 9, 31-37

Tarkhnishvili D., Gavashelishvili A., Murtskhvaladze M., (2017) Landscape Complexity in the Caucasus Impedes Genetic Assimilation of Human Populations More Effectively than Language or Ethnicity. Human Biology, vol: 88, 287-300

Tarkhnishvili D., Gabelaia M., (2017) Isolated population of the Middle Eastern Phoenicolacerta laevis from the Georgian Black Sea Coast, and its genetic closeness to populations from southern Turkey. Zoology in the Middle East, 1-5

Tarkhnishvili D., Murtskhvaladze M., (2017) Coincidence of genotypes at two loci in two parthenogenetic rock lizards: how backcrosses might trigger adaptive speciation. Biological Journal of the Linnean Society, vol: 121, 365–378

Kopaliani N., Ninua L., (2017) Records of anomalously white harbour porpoises and atypical pigmented short-beaked common dolphin in the Georgian Black Sea waters. J. Black Sea/Mediterranean Environment, vol: 23, 66-74

Japoshvili B., Mumladze L., (2017) The population of Carssius gibelio (Bloch, 1782) and its parasites in Madatapa Lake (South Georgia). Iranian Journal of Fisheries Sciences , 793-799

Javakhishvili Z., (1999) The Corncrake (Crex crex) in Georgia.

Okrostsvaridze A., (2017) Thorium Resources and their Energy Potential in Georgian Republic, the Caucasus. Energy Procedia, vol: 125, 291-299

Okrostsvaridze A., (2016) Pliocene-Quaternary Samtskhe-Javakheti Volcanic Highland, Lesser Caucasus – as a Result of Mantle Plumes Activity.

Akhalkatsi M., (2017) Alpine Crop Wild Relatives in Georgia (The Caucasus).

Akhalkatsi M., Nakhutsrishvili G., (2008) Habitats of Allium Species in Georgia.

Murvanidze M., Mumladze L., (2016) Oribatida diversity in different microhabitats of Mtirala National Park. The Acarological Society of Japan, vol: 25, 35-49

Murvanidze M., Mumladze L., (2016) Annotated checklist of Georgian oribatid mites. Zootaxa, vol: 4089,

Murvanidze M., Mumladze L., (2013) Oribatid mite colonization of sand and manganese tailing sites. ACAROLOGIA A quarterly journal of acarology, since 1959, vol: 53, 203-215

Murvanidze M., Mumladze L., (2015) Annotated List of Invertebrates of the Georgian Karst Caves. Georgian Academic Book, Tbilisi.

Murvanidze M., Mumladze L., (2014 ) Oribatid mite (Acari: Oribatida) diversity in different forest stands of Borjom-Kharagauli National Park (Georgia) . Persian Journal of Acarology, vol: 3, 257–276

Murvanidze M., Mumladze L., (2011) Landscape distribution of oribatid mites (Acari, Oribatida) in Kolkheti National Park (Georgia, Caucasus). Zoosymposia, vol: 6, 221–233

Mosulishvili M., (2017) A consolidated list of Triticum species and varieties of Georgia to promote repatriation of local diversity from foreign genebanks. Annals of Agrarian Science, vol: 15, 61-70

Akhalkatsi M., Mosulishvili M., (2003 ) Wild Orchids of Georgia.

Akhalkatsi M., Mosulishvili M., (2003 ) Diversity and conservation of Georgian orchids. Calamus, CGS Ltd, Tbilisi.

Akhalkatsi M., Mosulishvili M., (2004 ) Iris family in Georgia.

Akhalkatsi M., Mosulishvili M., (2004 ) Wetland orchid Spiranthes amoena in Georgia.

Akhalkatsi M., Mosulishvili M., (2004 ) Spiranthes amoena – a new species for flora of Georgia. Jour. Eur. Orch. , vol: 36, 745-754

Akhalkatsi M., Mosulishvili M., (2006) Orchids and their habitats in Georgia. Jour. Eur. Orch. , vol: 38,

Akhalkatsi M., Mosulishvili M., (2008 ) Conservation and Sustainable Utilization of Rare Medicinal Plants in Samtskhe-Javakheti..


Ekhvaia J., Akhalkatsi M., (2006) Comparative study of the quantitative parameters of berries and seeds in the autochthonous red grape varieties of the Kolkhis (Western Georgia). Proc. Georgian Acad. Sci., Biol. Ser. B, vol: 4, 38-46

Ekhvaia J., Akhalkatsi M., (2010) Morphological variation and relationships of Georgian populations of Vitis vinifera L. subsp. sylvestris (CC Gmel.) Hegi. Flora - Morphology, Distribution, Functional Ecology of Plants, vol: 205, 608-617

Ekhvaia J., Akhalkatsi M., (2014 ) Allele sizes in base pairs for SSR loci of local cultivars and wild grapevines of Georgia.

Barjadze S., Murvanidze M., Mumladze L., et al. (2015) Annotated List of the Invertebrates in the Caves of Racha Region (Western Georgia). Proceedings of the Institute of Zoology,

Bakhtadze N., Barjadze S., et al. (2010) Karyological Study of Three Species of the Genus Aphis (Hemiptera: Aphididae) from Georgia. BULLETIN OF THE GEORGIAN NATIONAL ACADEMY OF SCIENCES, vol: 4, 130-132

Bakhtadze N., Mumladze L., (2014) Karyological Studies in Terrestrial Mollusks (Mollusca: Gastropoda: Pulmonata) of Georgia.

Bakhtadze N., Mumladze L., (2015 ) Chromosome studies of some Georgian terrestrial molluscs (Mollusca: Gastropoda: Pulmonata).

Jorjiashvili N., Elashvili M., (2016) Seismic hazard analysis of Adjara region in Georgia. Natural Hazards, vol: 81, 745–758

Matcharashvili T., Chelidze T., Jorjiashvili N., et al. (2013) Discrimination between stochastic dynamics patterns of ambient noises (Case study for Oni seismic station). Acta Geophysica, vol: 61, 1-18

Matcharashvili T., Chelidze T., Jorjiashvili N., (2012) Scaling features of ambient noise at different levels of local seismic activity: A case study for the Oni seismic station. Acta Geophysica, vol: 50, 809-832

Gratiashvili N., Barjadze S., (2014 ) Morphometrics, life history and population biology of the Ponto-Caucasian slave-making ant Myrmoxenus tamarae (Hymenoptera: Formicidae). European Journal of Entomology, vol: 112 , 175-179

Gratiashvili N., Barjadze S., (2008 ) CHECKLIST OF THE ANTS (FORMICIDAE LATREILLE, 1809) OF GEORGIA. Proceedings of the Institute of Zoology , 130 -146

Nakhutsrishvili G., Akhalkatsi M., (2006) Biotope Types of the Treeline of the Central Greater Caucasus. Nature Conservation, Springer, Berlin, Heidelberg, 211-225

Tarkhnishvili D., (2000) Pre-Pleistocene Refugia and Differentiation between Populations of the Caucasian Salamander (Mertensiella caucasica). Molecular Phylogenetics and Evolution, vol: 14, 414-422

Sokhadze G., Cowgill E., et al. (2018) Active convergence between the Lesser and Greater Caucasus in Georgia: Constraints on the tectonic evolution of the Lesser–Greater Caucasus continental collision. Earth and Planetary Science Letters, vol: 481, 154–161

Bussmann R., Kikvidze Z., et al. (2016) Your Poison in My Pie—the Use of Potato (Solanum tuberosum L.) Leaves in Sakartvelo, Republic of Georgia, Caucasus, and Gollobordo, Eastern Albania. Economic Botany, vol: 70, 431–437

Kikvidze Z., (2001) Richness of Colchic vegetation: comparison between refugia of south-western and East Asia. BMC Ecology, 1-10

Nakhutsrishvili G., (2013) Georgia: Protected Areas. The Vegetation of Georgia (South Caucasus), Springer, Berlin, Heidelberg, 223-225

Okrostsvaridze A., (2011) The first data on industrial concentration of thorium and bismuth in hydrothermally altered Lower-Jurassic clay- shales of the Stori Canyon (southern slope of the Greater Caucasus, Kakheti). Bulletin on the Georgian national Academy of scienes, vol: 5,

Okrostsvaridze A., (2013) Phanerozoic continental crust evolution of the Inner Caucasian Microplate: The Dzirula massif.

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Gurchiani K., (2017) How Soviet is the Religious Revival in Georgia: Tactics in Everyday Religiosity. Europe-Asia Studies, vol: 69, 508-531

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Sukhishvili L., Javakhishvili Z., (2016) 3D Virtual Reality Applied in Tectonic Geomorphic Study of the Gombori Range of Greater Caucasus Mountains.

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Zein R., (2016) Happiness and Well-Being as Seen by Children and Young People in Georgia. GESJ: Education Science and Psychology, vol: 42, 3-13

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Nakhutsrishvili G., (2017) Plant Diversity of the Central Great Caucasus. Plant Diversity in the Central Great Caucasus: A Quantitative Assessment, Springer, Cham, 17-111

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Tushabramishvili N., (2013) Evolution, Cultural Influences or the Migrations in in Paleolithic of Georgia.

Tushabramishvili N., Sukhishvili L., (2012) Georgia on the Crossroad. Cultural Exchanges and Evidences for Different Distance Contacts in Middle and Upper Paleolithic. UISPP, 1-28

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Adler D., Tushabramishvili N., (2002) The latest Neandertals of the southern Caucasus: new dates and new data from Ortvale Klde, the Georgian Republic. Journal of Human Evolution, vol: 42, A2-A2

Adler D., Tushabramishvili N., (2009) Neanderthal life-ways in the Southern Caucasus.

Gabelaia M., Tarkhnishvili D., Murtskhvaladze M., (2015) Phylogeography and morphological variation in a narrowly distributed Caucasian rock lizard, Darevskia mixta. Amphibia-Reptilia, vol: 36,

Gabelaia M., Tarkhnishvili D., (2018) Use of three-dimensional geometric morphometrics for the identification of closely related species of Caucasian rock lizards (Lacertidae: Darevskia). Biological Journal of the Linnean Society, vol: 20, 1-9

Ninua L., Tarkhnishvili D., (2018) Phylogeography and taxonomic status of trout and salmon from the Ponto‐Caspian drainages, with inferences on European Brown Trout evolution and taxonomy. Ecology and Evolution, vol: 8, 1-14

Hughes N., Akhalkatsi M., (2009) Characterizing Betula Litwinowii Seedling Microsites at the Alpine-Treeline Ecotone, Central Greater Caucasus Mountains, Georgia. Arctic, Antarctic, and Alpine Research, vol: 41, 112-118