David Tarkhnishvili

Keywords Caucasus , Ethnogenesis , Glacial refugia , Human ecology , Landscape genetics , Language , Paternal lineage , Y-dna haplogroup ,

Country: Georgia

Organization:Ilia State University

Department:Institute of Ecology

ResearchGate profile: https://www.researchgate.net/profile/David_Tarkhnishvili

Google scholar profile: https://scholar.google.com/citations?user=EbTRHQYAAAAJ&hl=pt-BR

Academia.edu profile: https://iliauni.academia.edu/DavidTarkhnishvili

Facebook profile: https://www.facebook.com/david.tarkhnishvili?fref=ts

LinkedIn profile: https://www.linkedin.com/in/david-tarkhnishvili-a6176325



graduated from Tbilisi State University (Biology, zoology) (1982). PhD in Ecology from Research Inst. of Ecology, Ural Sci. Center at Sverdlovsk (pres. Ekaterinsburg), Russia, 1987. Research – interspecific competition of amphibians. Work experience – Tbilisi State University, Vietnam Tropical Center (1990), Bangor University, UK (1996-1997, post-doctoral fellow), Zoological Research Institute and Museum Al. Koenig, Bonn, Germany (1998; 1999-2003; post-doctoral fellow, researcher).Studies mostly in ecology, biogeography, population genetics and evolution of amphibians and reptiles. Over 100 published research papers. At Ilia State since 2006, employed as a dean of the school of Life Sciences, director of the Institute of Ecology, and dean of the School of Science and Engineering.

Research interests


  • Evolutionary biology
  • Speciation
  • Population ecology and genetics
  • Ecological modeling
  • Biogeography
  • Human population genetics
  • Caucasus
  • Amphibians and reptiles were the main research object for many years, however, in recent years the projects are on different organisms, including conifer plants, lizards, snails, domestic animals, and humans.


Tarkhnishvili D., (2012) Palaeoclimatic models help to understand current distribution of Caucasian forest species. Biological Journal of the Linnean Society, vol: 105, 231-248

Tarkhnishvili D., (2014) Human paternal lineages, languages, and environment in the caucasus. Human Biology, vol: 86, 113-130

Tarkhnishvili D., (2014) Historical biogeography of the caucasus. Nova Science Publishers, Inc., New York.

Tarkhnishvili D., (2008) Ecological divergence between two evolutionary lineages of the Caucasian salamander: Evidence from GIS analysis. Herpetological Journal, vol: 18, 155-163

Murtskhvaladze M., Gavashelishvili A., Tarkhnishvili D., (2010) Geographic and genetic boundaries of brown bear (Ursus arctos) population in the Caucasus. Molecular Ecology, vol: 19, 1829-1841

Sayim F., Tarkhnishvili D., (2009) Some water chemistry parameters of breeding habitats of the Caucasian salamander, Mertensiella caucasica in the Western Lesser Caucasus. Comptes Rendus - Biologies, vol: 332, 464-469

Tarkhnishvili D., (2001) Humid forest refugia, speciation and secondary introgression between evolutionary lineages: Differentiation in a Near Eastern brown frog, Rana macrocnemis. Biological Journal of the Linnean Society, vol: 74, 141 - 156

Tarkhnishvili D., (2016) Mitochondrial phylogeny of the darevskia saxicola complex: Two highly deviant evolutionary lineages from the easternmost part of the range. Herpetological Journal, vol: 26, 175-182

Tarkhnishvili D., Murtskhvaladze M., Gavashelishvili A., (2013) Speciation in Caucasian lizards: Climatic dissimilarity of the habitats is more important than isolation time. Biological Journal of the Linnean Society, vol: 109, 876-892

Tarkhnishvili D., Gavashelishvili A., et al. (2001) Land Use and Landscape Conservation in Southern Georgia.

Kopaliani N., Tarkhnishvili D., (2014) Gene Flow Between Wolf and Shepherd Dog Populations in Georgia (Caucasus). Journal of heredity, vol: 105, 345-353

Murtskhvaladze M., Kopaliani N., Tarkhnishvili D., (2012) Gene introgression between Gazella subgutturosa and G. marica: Limitations of maternal inheritance analysis for species identification with conservation purposes. Acta Theriologica, vol: 57, 383–386

Zazanashvili N., Tarkhnishvili D., (2016) CAUCASUS.

Akhalkatsi M., Tarkhnishvili D., (2012) Habitats of Georgia.

Hille A., Tarkhnishvili D., (2002) Morphometric, biochemical and molecular traits in Caucasian wood mice (Apodemus/Sylvaemus), with remarks on species divergence. Acta theriologica, vol: 47, 389-416

Gabelaia M., Tarkhnishvili D., (2017) Phylogenetic signals in scale shape in Caucasian rock lizards (Darevskia species). Zoologischer Anzeiger - A Journal of Comparative Zoology,

Tarkhnishvili D., Gavashelishvili A., Murtskhvaladze M., (2017) Landscape Complexity in the Caucasus Impedes Genetic Assimilation of Human Populations More Effectively than Language or Ethnicity. Human Biology, vol: 88, 287-300

Tarkhnishvili D., Gabelaia M., (2017) Isolated population of the Middle Eastern Phoenicolacerta laevis from the Georgian Black Sea Coast, and its genetic closeness to populations from southern Turkey. Zoology in the Middle East, 1-5

Tarkhnishvili D., Murtskhvaladze M., (2017) Coincidence of genotypes at two loci in two parthenogenetic rock lizards: how backcrosses might trigger adaptive speciation. Biological Journal of the Linnean Society, vol: 121, 365–378

Tarkhnishvili D., (2000) Pre-Pleistocene Refugia and Differentiation between Populations of the Caucasian Salamander (Mertensiella caucasica). Molecular Phylogenetics and Evolution, vol: 14, 414-422